• Klink, P.C.*, Teeuwen, R.R.*, Lorteije, J.A.M., & Roelfsema, P.R.R. (2022). Pop-in: the inversion of pop-out for a feature dimension during visual search in area V4 of the monkey cortex. bioRxiv (*Contributed equally).


  • PRIME-DRE Consortium* (2022). Toward next-generation primate neuroscience: A collaboration-based strategic plan for integrative neuroimaging. Neuron *Inc. Klink, P.C.
  • Klink, P.C., Chen, X., Vanduffel, W., & Roelfsema, P.R. (2021). Population receptive fields in non-human primates from whole-brain fMRI and large-scale neurophysiology in visual cortex. eLife, 10:e67304. pdf
    • Highlighted in: Ip, B., & Bridge, H. (2021). Magnetic Resonance: Mapping the visual world to the human brain. eLife, 10:e75171. pdf
  • Gau, R., Noble, S., Heuer, K., Bottenhorn, K.L., Bilgin, I.P., Yang, Y-F., Huntenburg, J.M., Bayer, J., Bethlehem, A.I., (...) , The Brainhack Community* (2021). Brainhack: developing a culture of open, inclusive, community-driven neuroscience. Neuron, S0896-6273(21)00231-2. *Inc. Klink, P.C. pdf
  • Klink, P.C., Aubry, J-F., Ferrera, V., Fox, A.S., Froudist-Walsh, S., Jarraya, B., Konofagou, E., Krauzlis, R., Messinger, A., Mitchell, A.S., Ortiz-Rios, M., Oya, H., Roberts, A.C., Roe, A.W., Rushworth, M.F.S., Sallet, J., Schmid, M.C., Schroeder, C.E., Tasserie, J., Tsao, D., Uhrig, L., Vanduffel, W., Wilke, M., Kagan, I., & Petkov, C.I. (2021). Combining Brain Perturbation and Neuroimaging in Non-human Primates. NeuroImage, 235, 118017. pdf
  • Messinger, A., Sirmpilatze, N., Heuer, K., Loh, K-K, Mars, R.B., Sein, J., Xu, T., Glen, D., Jung, B., Seidlitz, J., Taylor, P., Toro, R., Garza-Villarreal, E.A., Sponheim, C., Wang, X., Benn, R.A., Cagna, B., Dadarwal, R., Evrard, H.C., Garcia-Saldivar, P., Giavasis, S., Hartig, R., Lepage, C., Liu, C., Majka, P., Merchant, H., Milham, M.P., Rosa, M.G.P., Tasserie, J., Uhrig, L., Margulies, D.S., & Klink, P.C. (2021). A collaborative resource platform for non-human primate neuroimaging. NeuroImage, 226, 117519. pdf
  • Fox, A.s., Holley, D., Klink, P.C., Arbuckle, S.A., Barnes, C.A., Diedrichsen, J., Kwok, S.C., Kyle, C., Pruszynski, J.A., Seidlitz, J., Zhou, X-F, Poldrack, R.A., & Gorgolewski, K.J. (2021). Sharing voxelwise neuroimaging results from rhesus monkeys and other species with Neurovault. NeuroImage, 225, 117518. pdf
  • PRIME-DE Consortium* (2020). Accelerating the Evolution of Nonhuman Primate Neuroimaging. Neuron, 105(4), 600-603. *Inc. Klink, P.C. pdf
  • Milham M.P, Ai, L., Koo, B. (...), Klink, P.C., (...), Margulies D.S., & Schroeder, C.E. (2018) An Open Resource for Non-human Primate Imaging, Neuron, 100(1), 61-74.e2. pdf
  • Roelfsema, P.R., Denys, D., & Klink, P.C. (2018). Mind reading and writing: The future of neurotechnology. Trends in Cognitive Sciences, 2(7), 598–610. pdf
  • Klink, P.C., Boucherie, D., Denys, D., Roelfsema, P.R., & Self M. (2017). Interocularly merged face percepts eliminate binocular rivalry. Scientific Reports, 7(1), 7585. pdf
  • Klink, P.C., Jeurissen, D., Theeuwes, J., Denys, D., & Roelfsema, P.R. (2017). Working memory accuracy for multiple targets is driven by reward expectation and stimulus contrast with different time-courses. Scientific Reports, 7(1), 9082. pdf
  • Klink, P.C.*, Dagnino, B.*, Gariel-Mathis, M-A.*, & Roelfsema, P.R. (2017). Distinct feedforward and feedback effects of microstimulation in visual cortex reveals neural mechanisms of texture-segregation. Neuron. 95(1), 209–220.e3. (*Contributed equally) pdf
  • Chen, X., Possel, J.K., Wacongne, C., Van Ham, A., Klink, P.C., & Roelfsema, P.R. (2017). 3D printing and modeling of customized implants and surgical guides for non-human primates. Journal of Neuroscience Methods, 286, 38-55. pdf
  • Christophel, T.B.*, Klink, P.C.*, Spitzer, B., Roelfsema, P.R. & Haynes J-D (2017). A distributed account of working memory storage. Trends in Cognitive Sciences. 21(2), 111–124. (*Contributed equally) pdf
  • De Graaf, T.A., van Ee, R., Croonenberg, D., Klink, P.C., & Sack, A.T. (2017). Perceptual disappearance of a competition-free salient visual stimulus at the offset of binocular rivalry. Journal of Vision. 17(1):2, 1–18. pdf
  • Klink, P.C. & Roelfsema, P.R.R (2016). Binocular rivalry outside the scope of awareness. Proceedings of the National Academy of Sciences, 113(30), 8352–8354. pdf
  • Brascamp, J.W., & Klink, P.C., with a contribution of Levelt, W.J.A.M. (2015). The 'laws' of binocular rivalry: 50 years of Levelt's propositions. Vision Research. 109: 20-37. pdf
  • Klink, P.C., Jentgens, P., & Lorteije, J.A.M. (2014). Priority maps explain the roles of value, attention, and salience in goal-oriented behavior. Journal of Neuroscience. 34(42), 13867-13869. pdf
  • Klink, P.C., Oleksiak, A., Lankheet, M.J.M., & van Wezel, R.J.A. (2012). Intermittent stimulus presentation stabilizes neuronal responses in macaque area MT. Journal of Neurophysiology. 108, 2101-2114. pdf
  • Montijn, J.S., Klink, P.C., & van Wezel, R.J.A. (2012). Divisive normalization and neuronal oscillations in a single framework of selective visual attention. Frontiers in Neural Circuits. 6(22),1-38. pdf
  • Lankheet, M.J.M., Klink, P.C., & Noest, A.J. (2012). Spike-Interval Triggered Averaging Reveals a Quasi-Periodic Spiking Alternative for Stochastic Resonance in Catfish Electroreceptors. PLoS ONE 7(3): e32786. pdf
  • Klink, P.C., van Wezel, R.J.A., & van Ee, R. (2012). United we sense divided we fail: Neural mechanisms of context-driven perceptual disambiguation. Philosophical Transactions of the Royal Society B: Biological Sciences. 367(1591), 906-918. pdf
  • Oleksiak, A., Klink, P.C., Postma, A., van der Ham, I.J.M., Lankheet, M.J.M. & van Wezel, R.J.A. (2011). Spatial summation in macaque parietal area 7a follows a winner-take-all rule. Journal of Neurophysiology, 105(3), 1150-1158. pdf
  • Klink, P.C., Montijn, J.S., & van Wezel, R.J.A. (2010). Crossmodal duration perception involves perceptual grouping, temporal ventriloquism and variable internal clock rates. Attention, Perception & Psychophysics, 73(1), 219-236. pdf
  • Oleksiak, A., Postma, A., van der Ham, I.J.M., Klink, P.C., & van Wezel, R.J.A. (2010). A review of lateralization of spatial functioning in nonhuman primates. Brain Research Reviews. pdf
  • Klink, P.C., Brascamp, J.W., Blake, R., & van Wezel, R.J.A. (2010). Experience-driven plasticity in binocular vision. Current Biology, 20(16), 1464-1469. pdf
  • Klink, P.C., Noest, A.J., Holten, V., van den Berg, A.V., & van Wezel, R.J.A. (2009). Occlusion-related lateral connections stabilize kinetic depth through perceptual coupling. Journal of Vision, 9(10):20, 1-20. pdf
  • Klink, P.C., van Ee, R., & van Wezel, R.J.A. (2008). General validity of Levelt’s propositions reveals common computational mechanisms for visual rivalry. PLoS ONE, 3(10): e3473. pdf
  • Klink, P.C., Van Ee, R., Nijs, M.M., Brouwer, G.J., Noest, A.J., & van Wezel, R.J.A. (2008). Early interactions between neuronal adaptation and voluntary control determine perceptual choices in bistable vision. Journal of Vision, 8(5):16, 1-18. pdf
  • Klink, P.C. (2008). Some spikes are more informative than others. Journal of Neuroscience, 28(19), 4844-4845. pdf

Book chapters

  • Klink, P.C., Self, M.W., Lamme, V.A., & Roelfsema, P.R. (2015). Theories and methods in the scientific study of consciousness. The Constitution of Phenomenal Consciousness: Towards a Science and Theory, ed. S. Miller, Advances in Consciousness Research. John Benjamins Publishing Company.
  • Klink, P.C., van Wezel, R.J.A., & van Ee, R. (2014). The future of binocular rivalry research: Reaching through a window on consciousness. The Constitution of Visual Consciousness: Lessons from Binocular Rivalry, ed. S. Miller, Advances in Consciousness Research. John Benjamins Publishing Company.

PhD thesis

  • Klink, P.C. (2011). Neural mechanisms of context-driven conscious visual perception. Utrecht University. pdf

Selected abstracts

  • Klink, P.C., Chen, X., Vanduffel, W., Denys, D, & Roelfsema, P.R., (2019) Benchmarking population receptive field maps with fMRI and large-scale neurophysiological recordings in awake non-human primates. Society for Neuroscience, Annual Meeting 2016. Chicago.
  • Klink, P.C. (2018) Dawn of the bionic brains. CogniTalks, Amsterdam.
  • Klink, P.C., (2016) Top-down and bottom-up salience dynamically determine the precision of working memory encoding. Perception Day. Radboud University, Nijmegen.
  • Klink, P.C., (2016) Reward and salience determine the precision of working memory encoding with different time-courses. Society for Neuroscience, Annual Meeting 2016. San Diego.
  • Klink, P.C., (2016) Top-down priorities and bottom-up salience compete in macaque visual cortex. Amsterdam Vision Meeting.
  • Klink, P.C., Lorteije, J.A.M., van Vugt, B., & Roelfsema, P.R. (2015). Priority-related response modulations in macaque visual cortex. FENS Brain Conference 'Bridging Neural Mechanisms & Cognition'. Copenhagen.
  • Klink, P.C., Lorteije, J.A.M., van Vugt, B., & Roelfsema, P.R. (2015). Bottom-up and top-down priorities modulate responses in macaque visual cortex. Society for Neuroscience, Annual Meeting 2015. Chicago.
  • De Graaf, T.A., van Ee, R., Croonenberg, D., Klink, P.C., & Sack, A.T. (2015) Disrupting binocular rivalry can lead to lasting visual suppression: psychophysics and computational modeling. Human Brain Mapping 2015.
  • Arsenault, J.T., Rima, S., Klink, P.C., Stemmann, H., & Vanduffel, W. (2013) Electrical microstimulation of the ventral tegmental area reinforces cue selection in monkeys. Society for Neuroscience, Annual Meeting 2013.
  • Klink, P.C., (2012) Perceptual stabilization of ambiguous visual input: a synthesis of perception, computation and neurophysiology. ECVP 2012, Alghero, Italy.
  • Klink, P.C., Oleksiak, A., Postma, A., van der Ham, I.J.M., Lankheet, M.J.M., & van Wezel, R.J.A. (2011). A ‘matched filter’ mechanism of visual short-term memory in macaque parietal area 7a. Annual meeting of the Society for Neuroscience, Washington DC, USA.
  • Klink, P.C. (2011). Experience-driven plasticity in adult binocular vision. 9th Dutch EndoNeuroPsycho Meeting, Lunteren, the Netherlands.
  • Klink, P.C., Lankheet, M.J.M., & van Wezel, R.J.A. (2011). Intermittent motion stimuli stabilize neuronal responses in area MT: Implications for the perceptual stabilization of visual ambiguities. Vision Science Society Annual Meeting, Naples FL, USA.
  • van Wezel, R.J.A., Lankheet, M.J.M., & Klink, P.C. (2010). The time-course of perceptual stabilization induces response stabilization in macaque area MT neurons. Society for Neuroscience, 74.12.
  • Klink, P.C., Noest, A.J., & Lankheet, M.J.M. (2010). Spike-pair triggered averaging reveals a function for spike generation in neural coding. The Federation of European Neuroscience Societies Forum, Amsterdam, the Netherlands.
  • Klink, P.C., Brascamp, J.W., Blake, R., & Van Wezel, R.J.A. (2010). Plasticity of interocular inhibition with prolonged binocular rivalry. Vision Science Society Annual Meeting, Naples FL, USA.
  • Klink, P.C., Lankheet, M.J.M., & Van Wezel, R.J.A. (2009). The neural signature of stabilized ambiguous stucture-from-motion in macaque area MT. Society for Neuroscience, 558.19.
  • Klink, P.C. (2009). Perception of ambiguous figures in spatiotemporal context. 8th Dutch EndoNeuroPsycho Meeting, Doorwerth, the Netherlands.
  • Klink, P.C., Noest, A.J., & Van Wezel, R.J.A. (2009). Amodal spatial facilitation resolves local ambiguities of kinetic depth. Journal of Vision, 9(8):275, 275a.
  • Klink, P.C., & van Wezel, R.J.A. (2009). Area MT and the stabilized perception of bistable structure-from-motion. European Primate Neurobiology Meeting, Goettingen, Germany.
  • Klink, P.C., Van Ee, R., & Van Wezel, R.J.A. (2008). Levelt’s propositions generalized for bi-stable structure-from-motion: Common computational mechanisms of visual rivalry. Perception 37 ECVP Abstract Supplement, page 28.
  • Klink, P.C., Van Ee, R., Noest, A.J., & Van Wezel, R.J.A. (2007). Early interactions among perceptual history, stimulus timing and voluntary control determine perceptual decisions in bistable vision. Society for Neuroscience, 507.14.4.
  • Noest, A.J., Van Ee, R., Van Wezel, R.J.A., Klink, P.C., & Nijs, M.M. (2007). Bistable percept-choice dynamics explained by early interactions between stimulus timing, voluntary bias, and perceptual history. Perception 36 ECVP Abstract Supplement.


  • Klink, P.C. (2007). Attention vs. Contrast for the Single Neuron: Does the analogy hold? Journal of Neuroscience, eLetter. online